三年来，Betty从一个什么都不懂的小助理，生长为Mode编辑。从令Daniel烦厌的小卒，成长为 Daniel最得力的助手和最贴心的朋友。从一个自卑的小女孩，成长为有主见的职业女性。虽然Betty在他人眼中还只是一只丑小鸭，但凭着自信，聪明和坚韧，也终于迎来了出头天。
上一季中，Betty终于如愿以偿得到自己梦寐以求的编辑工作，但是令她意想不到前男友 Matt居然成为了新上司。Daniel事业迎来了新高峰，却失去了挚爱的Molly，Wilhelmina 试图寻找Connor的同时，一个神秘人的出现令她大吃一惊。 Marc没有得到编辑的职位，沮丧无比。这一切都为我们留下一连串的悬念。
踏上新的工作岗位，Betty能否胜任？又将怎样面对自己的新上司兼前男友Matt？Daniel 失去了挚爱的Molly又将何去何从？令Wilhelmina大吃一惊的神秘人是否Connor？他们又会有什么新阴谋？竞争上岗失败的Marc是安于做一名小助理，还是转战Elle成为Betty的对手？Amanda生父依然成谜，本季会否解开谜团呢?
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